List of Published Papers about Phorids by Dr. Brian V. Brown

Brown, B.V. 1986. Striking sexual dimorphism and its taxonomic consequences among New World phorid flies (Diptera). Proceedings of the Entomological Society of Washington. 88: 787-788.

Brown, B.V. 1987. Revision of the Gymnophora (Diptera: Phoridae) of the Holarctic Region: classification, reconstructed phylogeny and geographic history. Systematic Entomology. 12: 271-304.

Brown, B.V. 1987. Classification, reconstructed phylogeny and geographic history of the Neotropical phorid flies of the genus Gymnophora (Diptera: Phoridae). Journal of Natural History. 21: 1477-1524.

Brown, B.V. 1988. Additions to the phorid chapter in the Manual of Nearctic Diptera, Volume 2 (Diptera: Phoridae). The Canadian Entomologist. 120: 307-322.

Brown, B.V. 1989. Testing phylogenetic hypotheses: new species of Gymnophora (Diptera: Phoridae) from Japan and Nepal, with an updated world classification. Canadian Journal of Zoology. 67: 2543-2556.

Brown, B.V. 1990. Triphleba vitrinervis (Malloch), an unrecognized species of Crinophleba Borgmeier (Diptera: Phoridae). Proceedings of the Entomological Society of Washington. 92: 294-296.

Brown, B.V. 1990. New Nearctic Region records of Palearctic Megaselia species (Diptera: Phoridae). Quaestiones Entomologicae. 26: 701-702.

Brown, B.V. 1992. Life history, immature stages and undescribed male of Rhynchomicropteron (Diptera: Phoridae). Journal of Natural History. 26: 407-416.

Brown, B.V. 1992. Generic revision of Phoridae of the Nearctic Region and phylogenetic classification of Phoridae, Sciadoceridae and Ironomyiidae (Diptera: Phoridea). Memoirs of the Entomological Society of Canada. 164: 1-144.

Brown, B.V. 1993. Convergent adaptations in some Phoridae (Diptera) living in the nests of social insects: a review of the New World Aenigmatiinae, in G.E. Ball & H.V. Danks (editors). Systematics and entomology: diversity, distribution, adaptation and application. Memoirs of the Entomological Society of Canada. 165: 115-137.

Brown, B.V. 1993. Taxonomy and preliminary phylogeny of the parasitic genus Apocephalus, subgenus Mesophora (Diptera: Phoridae). Systematic Entomology. 18: 191-230. PDF

Brown, B.V. 1994. A replacement name for a species of Megaselia (Diptera: Phoridae). Proceedings of the Entomological Society of Washington. 96: 176.

Brown, B.V. 1994. Life history parameters and new host records of phorid parasitoids of fireflies. Coleopterists Bulletin. 48: 145-147. PDF

Brown, B.V. 1994. Descriptions and subfamily classification of some unusual Phoridae (Diptera). The Canadian Entomologist. 126: 703-707.

Brown, B.V. 1994. Another generic synonym resulting from sexually dimorphic Phoridae (Diptera). Proceedings of the Entomological Society of Washington. 96: 764.

Brown, B.V. 1994. Revision and new species of the Apocephalus (Mesophora) truncaticercus-infragroup (Diptera: Phoridae). Contributions in Science. No. 449, 7 pp. PDF

Brown, B.V. 1995. Review of the species of Anevrina Lioy (Diptera: Phoridae), with a new species and a revised world key. Entomological Problems. 25 (1994): 1-10. PDF

Brown, B.V. 1995. Response to Disney. Journal of Natural History. 29: 259-264.

Brown, B.V. 1995. Comments on the proposed conservation of Sphaerocera Latreille, 1804 and Borophaga Enderlein, 1924 (Insecta, Diptera). Bulletin of Zoological Nomenclature. 52: 182-183.

Brown, B.V. 1996. First record of the genus Plectanocnema Schmitz in North America (Diptera: Phoridae). Proceedings of the Entomological Society of Washington. 98: 608-609.

Brown, B.V. 1996. Preliminary analysis of a host shift: revision of the Neotropical species of Apocephalus, subgenus Mesophora (Diptera: Phoridae). Contributions in Science. No. 462, 36 pp. PDF

Brown, B.V. 1996. A further species of Apocephalus, subgenus Mesophora (Diptera: Phoridae) parasitic on stingless bees (Hymenoptera: Apidae: Meliponinae). Studia dipterologica. 3: 231-235.

Brown, B.V. 1997. Systematics and fossil evidence of host-parasitoid relationships of Calamiscus Borgmeier (Diptera: Phoridae). Journal of Natural History. 31: 1253-1259.

Brown, B.V. 1997. Parasitic phorid flies: a previously unrecognized cost to aggregation behavior of male stingless bees. Biotropica. 29: 370-372.

Brown, B.V. 1997. Revision of the Apocephalus attophilus-group of ant-decapitating flies (Diptera: Phoridae). Contributions in Science. No. 468, 60 pp. PDF

Brown, B.V. 1997. The undescribed male of Beckerina sinefurca Borgmeier and the recognition of Neotropical Beckerina (Diptera: Phoridae). Entomological Problems. 28: 155-158.

Brown, B.V. 1998. New species and records of Gymnophora Macquart (Diptera: Phoridae) from southeast Asia. Contributions in Science. No. 471, 13 pp. PDF

Brown, B.V. 1998. Auxanommatidia boreotis Beyer is a species of Veruanus (Diptera: Phoridae). Studia dipterologica. 5: 335-336.

Brown, B.V. 1999. Differential host use by neotropical phorid flies (Diptera: Phoridae) that are parasitoids of ants (Hymenoptera: Formicidae). Sociobiology. 33: 95-103.

Brown, B.V. 1999. Re-evaluation of the fossil Phoridae. Journal of Natural History. 33: 1561-1573.

Brown, B.V. 1999. Phorid flies of Costa Rican cloud forests. in Nadkarni, N.M. & Wheelwright, N.T. (editors). The Natural History and Ecology of Monteverde, Costa Rica: Background to Conservation. Oxford University Press.

Brown, B.V. 1999. A fossil Hypocera species (Diptera: Phoridae) and its phylogenetic implications. Studia dipterologica. 6: 273-277.

Brown, B.V. 2000. Revision of the "Apocephalus miricauda-group" of ant-parasitizing flies (Diptera: Phoridae). Contributions in Science. No. 482, 62 pp. PDF

Brown, B.V. 2000. Insects, overview. Pp. 479-484 in S.A. Levin (editor). Encyclopedia of Biodiversity, volume 3. Academic Press.

Brown, B.V. 2000. Five natural history notes on Neotropical phorid flies. Phorid Newsletter. 8: 1-2. PDF

Brown, B.V. 2001. Diversity of flies, gnats and mosquitoes. Pp. 815-826 in S.A. Levin (editor) Encyclopedia of Biodiversity, volume 2. Academic Press.

Brown, B.V. 2001. The species of Phora (Diptera: Phoridae) of Costa Rica. Revista de Biologia Tropical. 48 (2000): 977-982.

Brown, B.V. 2001. Taxonomic revision of Neodohrniphora, subgenus Eibesfeldtphora (Diptera: Phoridae). Insect Systematics and Evolution. 32: 393-409.

Brown, B.V. 2001. Beginning to study the bee-killing flies, the parasitoid genus Melaloncha Brues. Phorid Newsletter. 9: 1-5. PDF

Brown, B.V. 2002. A new primitive phorid (Diptera: Phoridae) from Baltic amber. Studia dipterologica. 8: 553-556.

Brown, B.V. 2002. Revision of the Apocephalus pergandei-group of ant-decapitating flies (Diptera: Phoridae). Contributions in Science. No. 496, 58 pp. PDF

Brown, B.V. 2003. A new fossil species of Triphleba (Diptera: Phoridae) with bifurcate antennae. Studia dipterologica. 10: 195-197.

Brown, B.V. 2003. In memory of Laurence W. Quate (1925-2002) [scientific obituary]. Studia Dipterologica. 10: 353-356.

Brown, B.V. 2004. Revision of the subgenus Udamochiras of Melaloncha bee-killing flies (Diptera: Phoridae: Metopininae). Zoological Journal of the Linnean Society. 140: 1-42. PDF

Brown, B.V. 2004. Revision of the Melaloncha cingulata-group of bee-killing flies (Diptera: Phoridae). Annals of the Entomological Society of America. 97: 386-392.

Brown, B.V. 2004. Diversity of ant-decapitating flies (Diptera: Phoridae: Apocephalus) from the ALAS project: new results and projections. Sociobiology. 44: 683-688.

Brown, B.V. 2005. A new Baltic amber Triphleba Rondani with greatly enlarged palpi (Diptera: Phoridae). Studia Dipterologica. 11: 549-552.

Brown, B.V. 2005. Revision of the Melaloncha furcata-group of bee-killing flies (Diptera: Phoridae). Insect Systematics and Evolution. 36: 241-258. PDF

Brown, B.V. 2005. Malaise trap catches and the crisis in Neotropical dipterology. American Entomologist. 51: 180-183. PDF

Brown, B.V. 2005. Classification of two poorly-known genera of African Phoridae (Diptera). African Invertebrates. 46:133-140. PDF

Brown, B.V. 2006. Revision of the untreated taxa of Melaloncha s. s. bee-killing flies (Diptera: Phoridae). Zootaxa. 1280: 68 pp. (PDF available upon request from the author)

Brown, B. V. 2007. A further new genus of primitive phorid fly (Diptera: Phoridae) from Baltic amber and its phylogenetic implications. Contributions in Science. 513: 1-14. PDF

Brown, B. V. 2007. The problematic Neotropical genus Cyrtophorina Borgmeier & Prado (Diptera: Phoridae). Journal of the Entomological Society of Ontario. 137: 117-130. PDF

Brown, B. V. 2007. Novel character states in fossil species of modern phorid genera (Diptera: Phoridae). Studia dipterologica. 14: 107-116. PDF

Brown, B. V. 2008. New records and a new species of the Dohrniphora longirostrata group (Diptera: Phoridae). Journal of the Kansas Entomological Society. 81: 204-207. PDF

Brown, B. V. 2009. A species of Xenotriphleba Buck (Diptera: Phoridae) from Baltic amber. Proceedings of the Entomological Society of Washington. 111: 33-37. PDF

Brown, B. V. 2009. The undescribed male of Ctenopleuriphora Liu (Diptera: Phoridae). Studia dipterologica. 15: 185-187. PDF

Brown, B.V. 2009. Three new species of parasitoid Phoridae (Diptera) from the Neotropical Region. Sociobiology. 54: 715-722. PDF

Brown, B.V. 2010. An enigmatic new genus and species of phorid fly from Baltic amber. Studia dipterologica. 16: 43-46. PDF

Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley, and M. Zumbado (editors). 2009. Manual of Central American Diptera, Volume 1. National Research Council Press, Ottawa, Canada. 714 pp. BOOKS PAGE

Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley, & M. Zumbado (editors). 2010. Manual of Central American Diptera, Volume 2. National Research Council Press, Ottawa, Canada. xvi + pp. 715-1442. BOOKS PAGE

Brown, B.V. and M. Buck. 1998. Review of the genus Hypocera Lioy (Diptera: Phoridae). Entomologica scandinavica. 29: 47-56.

Brown, B.V. and R.H.L. Disney. 2009. Validation of the name Diocophora multichaeta (Diptera: Phoridae). Proceedings of the Entomological Society of Washington. 111: 128-131. PDF

Brown, B.V., R.H.L. Disney, L. Elizalde, & P.J. Folgarait. 2010. New species and new records of Apocephalus Coquillett (Diptera: Phoridae) that parasitize ants (Hymenoptera: Formicidae) in South America. Sociobiology. 55: 165-190. PDF

Brown, B. V., L. F. Faust, & J. M. Cicero. 2009. A cantharid beetle host for a parasitic phorid fly new to the Nearctic Region (Coleoptera: Cantharidae; Diptera: Phoridae). Proceedings of the Entomological Society of Washington. 111: 898-899. PDF

Brown, B.V. and D.H. Feener, Jr. 1991. Life history parameters and description of the larva of Apocephalus paraponerae (Diptera: Phoridae), a parasitoid of the giant tropical ant Paraponera clavata (Hymenoptera: Formicidae). Journal of Natural History. 25: 221-321.

Brown, B.V. and D.H. Feener, Jr. 1991. Behavior and host location cues of Apocephalus paraponerae (Diptera: Phoridae), a parasitoid of the giant tropical ant, Paraponera clavata (Hymenoptera: Formicidae). Biotropica. 23: 182-187.

Brown, B.V. and D.H. Feener, Jr. 1993. Life history and immature stages of Rhyncophoromyia maculineura, an ant-parasitizing phorid fly (Diptera: Phoridae) from Peru. Journal of Natural History. 27: 429-434.

Brown, B.V. and D.H. Feener, Jr. 1995. Efficiency of two mass sampling methods for sampling phorid flies (Diptera: Phoridae) in a tropical biodiversity survey. Contributions in Science. No. 459, 10 pp. PDF

Brown, B.V. and D.H. Feener, Jr. 1998. Parasitic phorid flies (Diptera: Phoridae) associated with army ants (Hymenoptera: Formicidae: Ecitoninae, Dorylinae) and their conservation biology. Biotropica. 30: 482-487.

Brown, B. V., P. Folgarait, and L. Gilbert. 2003. A new species of Pseudacteon Coquillett (Diptera: Phoridae) attacking Solenopsis fire ants (Hymenoptera: Formicidae) in Argentina. Sociobiology. 41:685-688.

Brown, B. V., Francoeur, A. and R.L. Gibson. 1991. Review of the genus Styletta (Diptera: Phoridae), with description of a new genus. Entomologica scandinavica. 22: 241-250.

Brown, B. V. and G. Kung. 2004. Two new genera of Phoridae (Insecta: Diptera) from the Neotropical Region. Zootaxa. 554: 1-7. PDF

Brown, B.V. and G. Kung. 2006. Revision of the Melaloncha ungulata-group of bee-killing flies (Diptera: Phoridae). Contributions in Science. No. 507, 31 pp. PDF

Brown, B. V. and G. Kung. 2007. Revision of the New World Dohrniphora Dahl species with hind tibial setae (Diptera: Phoridae). Arthropod Systematics & Phylogeny. 65: 155-235. PDF

Brown, B.V. & G. Kung. 2010. Revision of the New World Dohrniphora Dahl species lacking large hind tibial setae (Diptera: Phoridae). Zootaxa. 2699: 1-142. (PDF available upon request from the author)

Brown, B. V. & E. G. LeBrun. 2010. Ant-decapitating flies, Apocephalus, subgenus Apocephalus Coquillett (Insecta: Diptera: Phoridae), of southern Arizona. Contributions in Science. 519: 1-24. PDF

Brown, B.V. and S.A. Marshall. 1984. Phorid flies (Diptera: Phoridae) associated with mushrooms in southern Ontario. Proceedings of the Entomological Society of Ontario. 115: 77-80.

Brown, B.V. and L. W. Morrison. 1999. New Pseudacteon (Diptera: Phoridae) from North America that parasitizes the native fire ant, Solenopsis geminata (Hymenoptera: Formicidae). Annals of the Entomological Society of America. 92: 308-311.

Brown, B. V. and H. Oliver. 2007. First records of Megaselia scalaris (Loew) and M. spiracularis Schmitz (Diptera: Phoridae) from New Zealand, with additional information on other worldwide species. New Zealand Entomologist. 30: 85-87. PDF

Brown, B. V. and H. Oliver. 2008. Two new genera of Phoridae (Diptera) from New Zealand. Zootaxa. 1933: 1-11. PDF

Brown, B. V. and H. Oliver. 2011. Male terminalia and undescribed female of Ceratoplatus fullerae Schmitz (Diptera: Phoridae) from New Zealand. New Zealand Entomologist. 34: 65-66. PDF

Brown, B.V. and E.M. Pike. 1990. Three new fossil phorid flies (Diptera: Phoridae) from Canadian Cretaceous amber. Canadian Journal of Earth Sciences. 27: 845-848.

Brown, B.V. and C.W. Sabrosky. 1994. Sphaerocera Latreille, 1804 and Borophaga Enderlein, 1924 (Insecta, Diptera): proposed conservation; Sphaerocera curvipes Latreille, 1805 and Phora flavimana Meigen, 1830: proposed conservation of the specific names. Bulletin of Zoological Nomenclature. 51: 312-315.

Brown, B. V., S. A. Schneider, & J. S. LaPolla. 2011. A new North American species of Pseudacteon (Diptera: Phoridae), parasitic on Nylanderia arenivaga (Hymenoptera: Formicidae). Annals of the Entomological Society of America. 104: 37-38. PDF

Brown, B. V. & P. T. Smith. 2010. The bee-killing flies, genus Melaloncha Brues (Diptera: Phoridae): a combined molecular and morphological phylogeny. Systematic Entomology. 35: 649-657. PDF

Corona, E.M. and B. V. Brown. 2004. Two new species of the Apocephalus attophilus-group of ant-decapitating flies (Diptera: Phoridae). Sociobiology. 44: 689-693.

Corona, E. M. and B.V. Brown. 2005. The Central American species of Diplonevra Lioy (Diptera: Phoridae). Zootaxa. 1050: 21-38. PDF

Disney, R.H.L. and B.V. Brown. 2003. A new species of Megaselia (Diptera: Phoridae) parasitising a Camponotus species (Hymenoptera: Formicidae) in Thailand. Sociobiology. 41: 403-410.

Disney, R. H. L. and B. V. Brown. 2009. Two additions to the list of phorid flies (Diptera: Phoridae) reported from the USA. Proceedings of the Entomological Society of Washington. 111: 539-540. PDF

Feener, D.H., Jr. and B.V. Brown. 1992. Reduced foraging of the tropical fire ant Solenopsis geminata (Hymenoptera: Formicidae), in the presence of parasitic phorid flies, Pseudacteon spp. Annals of the Entomological Society of America. 85: 80-84.

Feener, D.H., Jr. & B.V. Brown. 1993. Oviposition behavior of an ant-decapitating fly, Neodohrniphora curvinervis (Diptera: Phoridae) and defense behavior by its leaf-cutting ant host, Atta cephalotes (Hymenoptera: Formicidae). Journal of Insect Behavior. 6: 675-688.

Feener, D.H., Jr. & B.V. Brown. 1997. Diptera as parasitoids. Annual Review of Entomology. 42: 73-97.

Gonzalez, L. & B.V. Brown. 2004. New species and records of Melaloncha (Udamochiras) bee-killing flies (Diptera: Phoridae). Zootaxa. 730: 1-14. PDF

Gonzalez, V.H., B.V. Brown and M. Ospina. 2002. A new species of Megaselia (Diptera: Phoridae) associated with brood provisions of nests of Neocorynura (Hymenoptera: Halictidae). Journal of the Kansas Entomological Society. 75: 73-79.

Hall, D.W. and B.V. Brown. 1993. Pollination of Aristolochia littoralis (Aristolochiales: Aristolochiaceae) by males of Megaselia spp. (Diptera: Phoridae). Annals of the Entomological Society of America. 86: 609-613.

Hanson, P.S. and B.V. Brown. 1999. 4.6 Flies. Introduction. in Nadkarni, N.M. & Wheelwright, N.T. (editors). The Natural History and Ecology of Monteverde, Costa Rica: Background to Conservation. Oxford University Press.

Hibbs, P.J. and B.V. Brown. 1998. A new, extremely primitive species of Plethysmochaeta (Diptera: Phoridae) from South Africa. Studia dipterologica. 4 (1997): 367-369.

Kung, G. and B.V. Brown. 2000. The patterned-wing species of Coniceromyia (Diptera: Phoridae). Contributions in Science. No. 484, 10 pp. PDF

Kung, G. and B.V. Brown. 2000. Two new species of Coniceromyia (Diptera: Phoridae) from South America. Studia dipterologica. 7: 83-86.

Kung, G. and B.V. Brown. 2001. Two new patterned-wing species of Coniceromyia (Diptera: Phoridae) from Colombia. Studia dipterologica. 8: 267-270.

Kung, G. and B. V. Brown. 2004. Two new species of Megaselia Rondani (Diptera: Phoridae) from Costa Rica. Proceedings of the Entomological Society of Washington. 106: 751-756.

Kung, G. and B.V. Brown. 2005. New species of Dohrniphora related to D. longirostrata (Enderlein) (Diptera: Phoridae). Annals of the Entomological Society of America. 98: 55-62. PDF

Kung, G. And B.V. Brown. 2005. Three unusually divergent new genera of phorid flies (Diptera: Phoridae) from Costa Rica. Zootaxa. 1019: 43-52. PDF

Kung, G. and B. V. Brown. 2006. The Caribbean species of Dohrniphora Dahl (Diptera: Phoridae). Journal of Natural History. 40: 1931-1945. PDF

Marshall, S.A. and B.V. Brown. 1984. Sphaeroceridae (Diptera) associated with decaying fungi in Ontario. Proceedings of the Entomological Society of Ontario. 115: 43-48.

Morehead, S., J. Seger, D. H. Feener, Jr. and B. V. Brown. 2001. Evidence for a cryptic species complex in the ant parasitoid Apocephalus paraponerae (Diptera: Phoridae). Evolutionary Ecology Research. 3: 273-284.

Plowes, R. M., E. G. LeBrun, B. V. Brown, & L. E. Gilbert. 2009. A review of Pseudacteon (Diptera: Phoridae) that parasitize Solenopsis geminata complex (Hymenoptera: Formicidae). Annals of the Entomological Society of America. 102: 937-958. PDF

Quate, L. W. and B. V. Brown. 2004. Revision of the Neotropical Setomimini (Diptera: Psychodidae: Psychodinae). Contributions in Science. No. 500, 117 pp. PDF

Robinson, W. H. and B. V. Brown. 1993. Life history and immature stages of two species of Megaselia (Diptera: Phoridae) predatory on gall-inhabiting insects. Proceedings of the Entomological Society of Washington. 95: 404-411.

Seid, M.A. & B.V. Brown. 2009. A new host association of Commoptera solenopsidis (Diptera: Phoridae) with the ant Pheidole dentata (Hymnenoptera: Formicidae) and behavioral observations. Florida Entomologist. 92: 309-313. PDF

Smith, P. T. and B. V. Brown. 2006. Molecular evidence for the family classification of Tachinisca Kertész, 1903 (Diptera: Tephritoidea). Studia dipterologica. 13: 149-154.

Smith, P. T. and B. V. Brown. 2008. Utility of DNA sequences for inferring phylogenetic relationships and associating morphologically dissimilar males and females of the bee-killing flies, genus Melaloncha (Diptera: Phoridae). Annals of the Entomological Society of America. 101: 713-721.

Smith, P.T. & B.V. Brown. 2010. A molecular phylogenetic analysis of genus Anevrina (Diptera: Phoridae), with description of a new species, and updated world key. Zootaxa. 2397: 29-40. PDF