The Diptera, or true flies, are a huge group of insects that are familiar to most people. Worldwide there are about 153,000 validly described specie of flies, making them about 10% of the known diversity of life. Experts believe that most species of flies still have not been collected or described, however, and if all were known the group could number a million or more. Such an enormous number of species, coupled with their small body size, and the paucity of researchers working on them, makes Diptera one of the true frontiers of systematic research.

 

photo by S. A. Marshall

 
 
 

Flies are found worldwide on every continent, with few species living in Antarctica and with tremendous numbers in tropical regions. There is no analysis comparing the number of species of Diptera from different continents or biogeographical regions, but use of the existing Diptera catalogs (some of which are badly out of date) leads to the impression that the Palearctic (Old World temperate) Region is the area with the most species of flies. This is purely an artifact of research effort, however, with generations of European dipterists working on their local fauna while tropical regions remained virtually untouched. An informal survery of the some of the authors of chapters in the Manual of Central American Diptera reinforces the more logical conclusion that the New World tropics, including Central America, are the most species-rich areas of the world for flies.

 
 
 

Families of Diptera found in Central America and Tropical Mexico:

 
 

Nematocerous families:

 

Ptychopteridae
Culicidae
Dixidae
Corethrellidae
Chaoboridae
Ceratopogonidae
Chironomidae
Simuliidae
Blephariceridae
Bibionidae
Cecidomyiidae
Mycetophilidae
Ditomyiidae
Keroplatidae
Diadocidiidae
Lygistorrhinidae
Sciaridae
Tipulidae
Trichoceridae
Anisopodidae
Psychodidae
Scatopsidae

photo by S. A. Marshall

 
 
 

Lower Brachycera families:

 

Xylophagidae
Athericidae
Rhagionidae
Tabanidae
Vermileonidae
Pantophthalmidae
Stratiomyidae
Xylomyidae
Acroceridae
Nemestrinidae
Apioceridae
Asilidae
Bombyliidae
Mythicomyiidae
Mydidae
Scenopinidae
Therevidae
Dolichopodidae
Empididae

photo by S. A. Marshall

 
 
 

Lower Cyclorrhapha families:

 

Lonchopteridae
Platypezidae
Phoridae
Syrphidae
Pipunculidae

 

photo by S. A. Marshall

 

Schizophora families:

 

Micropezidae
Neriidae
Pseudopomyzidae
Tanypezidae
Somatiidae
Psilidae
Syringogastridae
Conopidae
Lonchaeidae
Ulidiidae
Platystomatidae
Tephritidae
Pyrgotidae
Ctenostylidae
Richardiidae
Piophilidae
Lauxaniidae
Chamaemyiidae
Paraleucopidae
Sciomyzidae
Ropalomeridae
Sepsidae
Clusiidae
Odiniidae
Agromyzidae
Anthomyzidae
Aulacigastridae
Periscelididae
Nannodastiidae
Asteiidae
Braulidae
Carnidae
Tethinidae
Canacidae
Coelopidae
Milichiidae
Cryptochaetidae
Chloropidae
Heleomyzidae
Chyromyidae
Sphaeroceridae
Curtonotidae
Camillidae
Drosophilidae
Diastatidae
Ephydridae
Inbiomyiidae

photo by S. A. Marshall

 
 
 

Calyptratae families:

 

Hippoboscidae
Streblidae
Nycteribiidae
Scathophagidae
Anthomyiidae
Fanniidae
Muscidae
Calliphoridae
Sarcophagidae
Rhinophoridae
Tachinidae
Oestridae

photo by S. A. Marshall